Richard B. Forward

Richard B. Forward

Research Professor

Bass Fellow

R. Forward works on the physiological ecology of marine crustaceans and fishes. Studies focus on sensory physiology and behavioral responses to environmental (e.g. light, temperature, salinity) and chemical cues, and biological rhythms. Recent studies have applied result to diel vertical migration and selective tidal stream transport.


Waite, H. R., K. G. Sanchez, and R. B. Forward. “Entrainment of the circadian rhythm in larval release of the crab Dyspanopeus sayi by temperature cycles.” Marine and Freshwater Behaviour and Physiology 50, no. 1 (January 2, 2017): 41–54.
Welch, J. M., K. A. Reinsel, E. E. Waddell, A. J. Wright, and R. B. Forward. “Chemical cues from adult fiddler crabs stimulate molting of conspecific megalopae: Evidence from field-caught individuals.” Marine Ecology Progress Series 553 (July 14, 2016): 147–53.
Cohen, J. H., and R. B. Forward. “Zooplankton Diel Vertical Migration - A Review of Proximate Control.” In Oceanography and Marine Biology: An Annual Review, 47:77–110, 2016.
Forward, Richard B., Kevin G. Sanchez, and Paul P. Riley. “Entrainment of the Circadian Rhythm in Egg Hatching of the Crab Dyspanopeus sayi by Chemical Cues from Ovigerous Females..” The Biological Bulletin 230, no. 1 (February 2016): 15–24.
Reinsel, K. A., J. M. Welch, A. O. Romero, E. M. Parks, K. M. McQueen, M. J. Smith, A. L. Price, et al. “Planktonic ingress of fiddler crab megalopae to the Newport River Estuary, NC: Evidence for semilunar periodicity in Uca pugnax and species-specific sampling bias by hog's hair larval collectors.” Marine Ecology Progress Series 523 (January 1, 2015): 105–13.

Recent Grants


BIOLOGY 493A: Research Independent Study (BIOLOGY 493A: Research Independent Study)

area(s) of expertise

Marine Science

Contact Information

135 Duke Marine Lab Rd
Beaufort, NC 28516


Ph.D., University of California at Santa Barbara (1969)
B.A., Stanford University (1965)