Richard B. Forward

Richard B. Forward

Research Professor

Bass Fellow

R. Forward works on the physiological ecology of marine crustaceans and fishes. Studies focus on sensory physiology and behavioral responses to environmental (e.g. light, temperature, salinity) and chemical cues, and biological rhythms. Recent studies have applied result to diel vertical migration and selective tidal stream transport.


Waite, HR, Sanchez, KG, and Forward, RB. "Entrainment of the circadian rhythm in larval release of the crab Dyspanopeus sayi by temperature cycles." Marine and Freshwater Behaviour and Physiology 50, no. 1 (January 2, 2017): 41-54.
Forward, RB, Sanchez, KG, and Riley, PP. "Entrainment of the Circadian Rhythm in Egg Hatching of the Crab Dyspanopeus sayi by Chemical Cues from Ovigerous Females." The Biological bulletin 230, no. 1 (February 2016): 15-24.
Reinsel, K, Welch, J, Romero, A, Parks, E, McQueen, K, Smith, M, Price, A, Clark, H, Zmina, S, Williams, C, and Forward, R. "Planktonic ingress of fiddler crab megalopae to the Newport River Estuary, NC: evidence for semilunar periodicity in Uca pugnax and species-specific sampling bias by hog’s hair larval collectors." Marine Ecology Progress Series 523 (March 16, 2015): 105-113.
Welch, J, Reinsel, K, Battles, K, Romero, A, Blaine, J, Sendi, R, and Forward, R. "Settlement of fiddler crab megalopae on a North Carolina (USA) sandflat: species identification using multiplex PCR provides evidence for selective settlement." Marine Ecology Progress Series 523 (March 16, 2015): 115-123.
Forward, RB, Moeller, BP, and Cohen, JH. "Circadian rhythm in larval release by the crab Rhithropanopeus harrisii: entrainment model." The Biological bulletin 226, no. 2 (April 2014): 92-101.

Recent Grants


BIOLOGY 792A: Research (BIOLOGY 792A: Research)
BIOLOGY 791TA: Tutorial (BIOLOGY 791TA: Tutorial)
BIOLOGY 493A: Research Independent Study (BIOLOGY 493A: Research Independent Study)

area(s) of expertise

Marine Science

Contact Information

135 Duke Marine Lab Rd
Beaufort, NC 28516


Ph.D., University of California at Santa Barbara (1969)
B.A., Stanford University (1965)