Richard B. Forward

Richard B. Forward

Research Professor

Bass Fellow


Forward, RB, Sanchez, KG, and Riley, PP. "Entrainment of the Circadian Rhythm in Egg Hatching of the Crab Dyspanopeus sayi by Chemical Cues from Ovigerous Females." The Biological bulletin 230, no. 1 (February 2016): 15-24.
Reinsel, KA, Welch, JM, Romero, AO, Parks, EM, McQueen, KM, Smith, MJ, Price, AL, Clark, HR, Zmina, SE, Williams, CA, and Forward, RB. "Planktonic ingress of fiddler crab megalopae to the Newport River Estuary, NC: evidence for semilunar periodicity in Uca pugnax and species-specific sampling bias by hog’s hair larval collectors." Marine Ecology Progress Series 523 (March 16, 2015): 105-113.
Welch, JM, Reinsel, KA, Battles, KA, Romero, AO, Blaine, JM, Sendi, RL, and Forward, RB. "Settlement of fiddler crab megalopae on a North Carolina (USA) sandflat: species identification using multiplex PCR provides evidence for selective settlement." Marine Ecology Progress Series 523 (March 16, 2015): 115-123.
Forward, RB, Moeller, BP, and Cohen, JH. "Circadian rhythm in larval release by the crab Rhithropanopeus harrisii: entrainment model." The Biological bulletin 226, no. 2 (April 2014): 92-101.
Ramus, AP, and Forward, RB. "The physiological ecology of the supratidal amphipod Talorchestia longicornis." Comp Biochem Physiol A Mol Integr Physiol 161, no. 2 (February 2012): 159-165.


BIOLOGY 792A: Research (BIOLOGY 792A: Research)
BIOLOGY 293A: Research Independent Study (BIOLOGY 293A: Research Independent Study)
ENVIRON 593: Independent Studies and Projects (ENVIRON 593: Independent Studies and Projects)
BIOLOGY 490T-1: Tutorial (BIOLOGY 490T-1: Tutorial)
BIOLOGY 791TA: Tutorial (BIOLOGY 791TA: Tutorial)

area(s) of expertise

Marine Science

Contact Information

135 Duke Marine Lab Rd
Beaufort, NC 28516


Ph.D., University of California at Santa Barbara (1969)
B.A., Stanford University (1965)